Зоологический журнал, 2024, № 10

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том 103       № 10       2024       Октябрь
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СОДЕРЖАНИЕ
Том 103, Номер 10, 2024
New species and new records of oribatid mites (Acari, Oribatida) from Villa Clara, Cuba     
S. G. Ermilov, A. V. Tolstikov	
3
Moraria (Baikalomoraria) okunevae sp. n. и Moraria (Baikalomoraria) ornofurcata sp. n.  
(Copepoda, Harpacticoida, Canthocamptidae) из озера Байкал
Т. М. Алексеева	
12
Половой аппарат самок видов рода Hololepta (Coleoptera, Histeridae,  
Histerinae, Hololeptini) и его значение для систематики группы     
В. В. Бичевой	
30
Холодостойкость и условия зимовки гусениц сибирского коконопряда  
(Dendrolimus sibiricus, Lepidoptera, Lasiocampidae) центрально-якутской популяции
Н. А. Булахова, Е. Н. Мещерякова, А. П. Бурнашева,  
Ю. В. Ермакова, Н. Н. Винокуров, Д. И. Берман	
55
Первая встреча биссы (Eretmochelys imbricata) и другие находки 
морских черепах на российском Дальнем Востоке
К. Д. Мильто, И. В. Маслова, С. Ю. Стефанов	
72
Случай смены партнера в гнездовой период у белого аиста  
(Ciconia ciconia L. 1758, Ciconiiformes, Ciconiidae) и его последствия
В. В. Павлова	
80
Перспективы сохранения амурского тигра (Panthera tigris altaica)  
на примере группировки в Хабаровском крае
В. С. Лукаревский, А. Ю. Олейников, С. А. Колчин	
88
Концентрация кортизола в волосах байкальских нерп  
(Pusa sibirica Gmelin 1788, Carnivora, Phocidae):  
связь с полом, возрастом животных и динамикой разрушения льда
П. Ю. Шибанова, М. А. Соловьёва, П. С. Журавлёва,  
Д. М. Глазов, В. В. Рожнов, С. В. Найденко	
107
Потери науки
Владислав Вильгельмович Хлебович (27.02.1932–22.02.2024)
И. С. Плотников, А. В. Смирнов, А. О. Смуров, Н. В. Аладин	
116

Contents
Volume 103, № 10, 2024
New species and new records of oribatid mites (Acari, Oribatida) from Villa Clara, Cuba
S. G. Ermilov, A. V. Tolstikov	
3
Moraria (Baikalomoraria) okunevae sp. n. and Moraria (Baikalomoraria) ornofurcata sp. n.  
(Copepoda, Harpacticoida, Canthocamptidae) from lake Baikal
T. M. Alekseeva	
12
The female reproductive system of the genus Hololepta (Coleoptera,  
Histeridae, Histerinae, Hololeptini) and its significance for the systematics of the group
V. V. Bichevoy	
30
Cold hardiness and overwintering conditions of larvae of the Siberian moth,  
Dendrolimus sibiricus (Lepidoptera, Lasiocampidae) from central Yakutia’s population
N. A. Bulakhova, E. N. Meshcheryakova, A. P. Burnasheva,  
Yu. V. Ermakova, N. N. Vinokurov, D. I. Berman	
55
The Hawksbill Turtle (Eretmochelys imbricata) in the Russian Far East  
and other new sea turtle records
K. D. Milto, I. V. Maslova, S. Yu. Stefanov	
72
A case of partner change in a pair during the nesting period  
in the White stork (Ciconia ciconia L. 1758, Ciconiiformes, Ciconiidae) and its consequences
V. V. Pavlova	
80
Perspectives of Amur tiger (Panthera tigris altaica) conservation,  
a population in the Khabarovsk Krai taken as an example
V. S. Lukarevskiy, A. Yu. Oleynikov, S. A. Kolchin	
88
Cortisol concentration in hair of the Baikal seal  
(Pusa sibirica, Gmelin, 1788, Carnivora, Phocidae): 
the effects of sex, age, and the dynamics of ice destruction
P. Yu. Shibanova, M. A. Solovyeva, P. S. Zhuravleva,  
D. M. Glazov, V. V. Rozhnov, S. V. Naidenko	
107
Obituary
Vladislav Vilgelmovich Khlebovich (27.02.1932–22.02.2024)
I. S. Plotnikov, A. V. Smirnov, A. O. Smurov, N. V. Aladin	
116

ЗООЛОГИЧЕСКИЙ ЖУРНАЛ,  2024, Том 103, № 10,  с.  3–11
This work is part of our faunistic and taxonomic 
study of oribatid mites (Acari, Oribatida) of Cuba (e. g. 
Ermilov et al., 2016, 2023a). The specimens presented 
in this paper were collected from the Villa Clara Province (located in the central part of the country). The 
main goals of our paper are: to present a list of all identified oribatid mites, and to describe two new species 
belonging to Muliercula Coetzer 1968 (Scheloribatidae) 
and Galumna (Galumna) Heyden 1826 (Galumnidae).
The genus 
was proposed by Coetzer 
(1968), with Muliercula muliercula Coetzer 1968
as the 
type species. At present, the genus comprises 13 species, 

and Oriental regions (Subías, 2022, online version 2023; 
Ermilov et al., 2023a). A revised generic diagnosis and 
an identification key to the known Muliercula species 
were provided by Ermilov and Rybalov (2023).

Galumna was proposed by Heyden (1826), 

Notaspis alatus Hermann, 1804 as the type species. 

species, which have a cosmopolitan distribution (Subías, 2022, online version 2023; Ermilov et al., 2023). 
Revised generic and subgeneric diagnoses, as well as 
an identification key to the Neotropical Galumna (Ga
lumna) species were provided by Ermilov and Klimov 
(2017) and Ermilov and Starý (2021), respectively.

Muliercula
seven Galumna (Galumna) species were recorded from 
Cuba (Jeleva et al., 1984; Ermilov, Tolstikov, 2015, 
2024; Ermilov et al., 2016, 2023, 2023a): M. spora 
Coetzer 1968; M. curvilineata Ermilov et Kontschán 
2023; G.
G.) angularis
G. G. arrugata Jeleva, Scull et Cruz 1984; G. (G.) australis (Berlese 1914); G. (G.) cubana Jeleva, Scull et Cruz 
1984; G. (G.) flabellifera Hammer 1958; G. (G.) gigantea 
Ermilov, Kolesnikov, Kontschán et Klimov 2023; and 
G. (G.) lunaris Jeleva, Scull et Cruz 1984.
METHODS
O b s e r v a t i o n  a n d  d o c u m e n t a t i o n. For 
measurement and illustration, specimens were mounted in lactic acid on temporary cavity slides. All measurements are in micrometers (μm). Body length was 
measured in lateral view, from the tip of the rostrum to 
the posterior edge of the notogaster. Notogastral width 
refers to the maximum width in dorsal aspect (behind 
pteromorphs). Setal lengths were measured perpendicular to their long axes, accounting for curvature. Formulas for leg solenidia are given in square brackets according to the sequence genu-tibia--tarsus. Drawings 
were made with a camera lucida using an “Olympus 
CX43RF” transmission light microscope.
T e r m i n o l o g y. Morphological terminology used 
in this paper mostly follows that used in the papers on 
Muliercula and Galumna (e. g., Grandjean, 1936, 1956; 
Ermilov, Rybalov, 2023; Ermilov et al., 2023, 2023a); 
also, see Norton (1977) for leg setal nomenclature, and 
Norton and Behan--Pelletier (2009) and Ermilov and 
Klimov (2017) for overview.
УДК 595.4
NEW SPECIES AND NEW RECORDS OF ORIBATID MITES 
(ACARI, ORIBATIDA) FROM VILLA CLARA, CUBA
© 2024 S. G. Ermilov*, A. V. Tolstikov**
Tyumen State University, X-BIO Institute, Tyumen, 625003 Russia
*e-mail: ermilovacari@yandex.ru
**e-mail: atolus@yahoo.com
Received December 20, 2023
Revised October 01, 2024
Accepted February 09, 2024
The present study is based on the oribatid mite material collected from the Villa Clara Province, Cuba. 
A list of 17 species, belonging to 11 genera and nine families, is presented; of them, one genus and three 
species are reported from Cuba for the first time. Two new species – Muliercula ornamenta Ermilov sp. n. 
(Scheloribatidae) and Galumna (Galumna) villaclaraensis Ermilov sp. n. (Galumnidae) – are described 
from soil litter with decaying leaves.
Keywords: Muliercula, Galumna, morphology, taxonomy, Neotropical fauna
DOI: 10.31857/S0044513424100017, EDN: tnburx

ERMILOV, TOLSTIKOV
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
Том 103
№ 10
2024
A b b r e v i a t i o n s. Prodorsum: lam = lamella; 
tlam = translamella; slam = sublamella; Al = sublamellar porose area; tu = tutorium; L = lamellar line; S 
= sublamellar line; N = prodorsal leg niche; E, T = lateral ridges of body; ro, le, in, bs, ex = rostral, lamellar, 
interlamellar, bothridial, and exobothridial setae, respectively; Ad = dorsosejugal porose area; D = dorsophragma; P = pleurophragma. Notogaster: c, la, lm, lp, 
h, p = notogastral setae; Aa, A1, A2, A3 = porose areas; 
Sa, S1, S2, S3 = saccules; mp = median pore; ia, im, ip, 
ih, ips = lyrifissures; gla = opisthonotal gland opening. 
Gnathosoma: a, m, h = subcapitular setae; or = adoral 
seta; as = axillary saccule. Epimeral and lateral podosomal regions: 1a, 1b, 1c, 2a, 3a, 3b, 3c, 4a, 4b = epimeral 
setae; z = aperture of supracoxal gland; PdI, PdII = pedotecta I and II, respectively; dis = discidium; cir = 
circumpedal carina. Anogenital region: g, ag, an, ad = 
genital, aggenital, anal, and adanal setae, respectively; 
iad = adanal lyrifissure; Amar = marginal porose area; 
po = preanal organ. Legs: Tr, Fe, Ge, Ti, Ta = trochanter, femur, genu, tibia, and tarsus, respectively; pa = porose area; ω, σ, φ = solenidia; ɛ = famulus; d, l, v, ev, 
bv, ft, tc, it, p, u, a, s, pv, pl = setae.
LIST OF IDENTIFIED TAXA
Locality: Cuba, Villa Clara Province, Baños de Elguea, soil litter with decaying leaves in a pasture under 
a tree (unknown data and collector; collection of the 
Tyumen State University Museum of Zoology, Tyumen, 
Russia). Distribution: mostly from Subías (2022, online 
version 2023). Ptyctimous mites not included.
Nothridae
Nothrus anauniensis Canestrini et Fanzago 1877: 1 
ex. Distribution: Cosmopolitan. New record of the species in Cuba.
Otocepheidae
Pseudotocepheus sexidimorphus (Călugăr et Vasiliu 
1977): 5 ex. Distribution: Cuba. R e m a r k s. Our specimens are similar to those from the original description 
(Vasiliu, Călugăr, 1977) and redescription (Ermilov, 
2020), but they have (versus do not have) one additional 
prodorsal tubercle between medial prodorsal condyles. 
We believe that there is morphological variability.
Carabodidae
Kalloia simpliseta Mahunka 1985: 15 ex. Distribution: Neotropical, Côte d’Ivoire.
Tectocepheidae
Tectocepheus velatus (Michael 1880): 1 ex. Cosmopolitan.
Mochlozetidae
Mochlozetes penetrabilis Grandjean 1930: 7 ex. Distribution: Tropical, Japan.
Oribatulidae
Spinoppia magniserrata Higgins et Woolley 1966: 
1 ex. Distribution: Neotropical. New record of the genus and species in Cuba.
Haplozetidae
Rostrozetes ovulum (Berlese 1908): 1 ex. Distribution: 
Tropical, Subtropical.
Scheloribatidae
Muliercula ornamenta Ermilov sp. n.: 16 ex.
Scheloribates fimbriatus Thor 1930: 13 ex. Distribution: Cosmopolitan.
Scheloribates praeincisus praeincisus (Berlese 1910): 9 
ex. Distribution: Tropical, southern Holarctic.
Galumnidae
Galumna (Galumna) flabellifera Hammer 1958: 1 ex. 
Distribution: Tropical, Subtropical.
Galumna (Galumna) lunaris Jeleva, Scull et Cruz 1984: 
2 ex. Distribution: Cuba.
Galumna (Galumna) villaclaraensis Ermilov sp. n.: 2 ex.
Galumna (Galumna) sp.: 1 ex.
Galumna (Neogalumna) araujoi (Pérez--Íñigo et Baggio 
1994): 83 ex. Distribution: Neotropical. New record of the 
species in Cuba. 
Pergalumna brasiliensis (Sellnick 1923): 86 ex. Distribution: Neotropical. New record of the species in Cuba. R e 
m a r k s. Our specimens are similar to those from the original description (Sellnick, 1923), but they have oval (versus 
oblong) notogastral porose area A1. We believe that there is 
morphological variability.
Pergalumna silvatica Hammer 1961: 16 ex. Distribution: 
Neotropical.
The list includes 17 species belonging to 11 genera and 
nine families. One genus (Spinoppia) and three species (Nothrus anauniensis, Galumna (Neogalumna) araujoi, Pergalumna brasiliensis) are recorded for the first time from Cuba. 
Of the 14 species (excluding two new species and one unidentified species), two are registered only in Cuba, four are 
known from the Neotropical region, eight have a broader 
distribution (more than one geographical region).
TAXONOMY
Muliercula ornamenta Ermilov sp. n.
(Figs 1–2)
T y p e  m a t e r i a l. Holotype (♂) and 15 paratypes 
(9♂♂, 6♀♀): Cuba, Villa Clara Province, Baños de Elguea, 

	
NEW SPECIES AND NEW RECORDS OF ORIBATID MITES (ACARI, ORIBATIDA)
5
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
Том 103
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2024
soil litter with decaying leaves in a pasture under a tree 
(unknown data and collector; collection of the Tyumen 
State University Museum of Zoology, Tyumen, Russia).
The holotype is deposited in the collection of the 
Senckenberg Museum of Natural History, Görlitz, Germany; 15 paratypes are deposited in the collection of 
the Tyumen State University Museum of Zoology, Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.
D i a g n o s i s o f  a d u l t. Body length: 555–675. 
Prodorsum sparsely foveolate; notogaster striate, some 
striae in anterior notogastral part forming reticulate pattern; epimeral and anogenital regions sparsely lineolate 
and foveolate. Rostrum protruding, narrowly rounded. 
Translamella interrupted medially; lateral keel-shaped 
ridge and prolamella absent. Rostral, lamellar and interlamellar setae setiform, barbed; in > le > ro; rostral 
seta inserted dorsally; bothridial seta clavate, barbed. 
All setae short, needleform. Epimeral formula: 3–1–
3–2. Discidium and circumpedal carina well developed. 
Heterotridactylous.
D e s c r i p t i o n  o f  a d u l t. Measurements. Body 
length: 615 (holotype), 555–630 (male paratypes),  
Fig. 1. Muliercula ornamenta Ermilov sp. n., adult (gnathosoma and legs not shown): a – dorsal view, b – ventral view, c – 
right lateral view, d – posterior view. Scale bar 200 µm.

ERMILOV, TOLSTIKOV
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
Том 103
№ 10
2024
630–675 (female paratypes); body width: 405 (holotype), 405–435 (male paratypes), 465–510 (female 
paratypes).
Integument. Body color brown. Lateral side of body 
partially with densely microgranulate cerotegument. 
Body surface with microgranulate sculpturing (visible under high magnification, ×1000); prodorsum 
with small, sparse foveolae; notogaster striate but striae in anterior notogastral part forming reticulate pattern; epimeral and anogenital regions sparsely lineolate 
(short, depressed lines) and with small, sparse foveolae; 
subcapitular mentum, leg femora I–IV and trochanters 
III, IV sparsely lineolate.
Prodorsum. Rostrum protruding, narrowly rounded (nearly pointed in two paratypes). Lamella about 
1/2 length of prodorsum; translamella linear, interrupted medially; lateral keel-shaped ridge and prolamella absent; tutorium well developed, comparatively short; sublamella poorly visible. Sublamellar porose area (11– 13 × 7) oval. Rostral (64–67), lamellar 
(112– 127), and interlamellar (161–180) setae setiform, nearly straight, barbed; rostral seta inserted 
Fig. 2. Muliercula ornamenta Ermilov sp. n., adult: a – subcapitulum, ventral view; b – leg I, left, paraxial view; c – leg II 
(without tarsus), right, antiaxial view; d – leg III (without tarsus), left, antiaxial view; e – leg IV, left, antiaxial view. Scale 
bars 50 µm.

	
NEW SPECIES AND NEW RECORDS OF ORIBATID MITES (ACARI, ORIBATIDA)
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ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
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dorsally; exobothridial seta (49–56) setiform, thin, 
slightly barbed; bothridial seta (90–94) clavate, barbed; 
length of bothridial stalk approximately equal to head. 
Dorsosejugal porose area not observed.
Notogaster. Anterior notogastral margin convex medially. Pteromorph broadly rounded laterally. All setae 
(6) needleform. Four pairs of saccules drop-like. Opisthonotal gland opening and all lyrifissures distinct.
Gnathosoma. Generally, similar to Muliercula 
(e. g. Ermilov et al., 2023a). Subcapitulum size: 
139– 146 × 112– 116; subcapitular setae (a, m: 22–26; 
h: 35–37) setiform, slightly barbed; m thinnest; both 
adoral setae (13–15) setiform, barbed. Palp length: 
79– 86; postpalpal seta (6) spiniform, smooth. Chelicera 
length: 142– 146; setae (cha: 43–45; chb: 28–30) 
setiform, barbed.
Epimeral region. Epimeral formula: 3–1–3–2; setae 
(1a, 2a, 3a, 4a, 4b: 22–26; 1b, 1c, 3b, 3c: 34–37) setiform, thin, slightly barbed; 4a, 4b thinnest. Discidium 
and circumpedal carina well developed.
Anogenital region. Genital (17–19), aggenital 
(17– 19), anal (19–22), and adanal (22–26) setae setiform, thin, slightly barbed. Adanal lyrifissure distinct. 
Marginal porose area present, complete.
Legs. Heterotridactylous; all claws barbed on dorsal 
side; both lateral claws with small tooth distoventrally. Dorsoparaxial porose area on femora I–IV and on 
trochanters III, IV, ventrodistal porose area on tibia I–
IV and proximoventral porose area on tarsi I–IV present. Formulas of leg setation and solenidia: I (1– 5–3–
4– 20) [1–2–2], II (1–5–2–4–15) [1–1–2], III 
(2– 3–1–3–15) [1–1–0], IV (1–2–2–3–12) [0–1–0]; 
homology of setae and solenidia indicated in Table 1. 
Solenidia ω1 on tarsus I, ω1 and ω2 on tarsus II, and σ 
on genu III thickened, rounded distally; other solenidia 
setiform or subflagellate.
C o m p a r i s o n. Muliercula ornamenta Ermilov sp. n. is similar to M. curvilineata Ermilov and 
Kontschán 2023 from Cuba (see Ermilov et al., 2023a) 
in the presence of striate notogaster. However, the new 
species differs from the latter in the body ornamentation 
(prodorsum foveolate, anterior part of notogaster with 
reticulate pattern, ventral plate lineolate and foveolate 
versus prodorsum not foveolate, anterior part of notogaster not reticulate, ventral plate striate).
Also, M. ornamenta Ermilov sp. n. is similar to Fijibates dlouhyi (Mahunka 1984) from the Neotropical 
region and India (see Mahunka, 1984) in the presence 
of striate notogaster, with some striae forming reticulate pattern in anterior part of notogaster. However, 
the new species differs from F. dlouhyi in the following 
morphological traits: narrow (versus broad) lamella; absence (versus presence) of prolamella; presence (versus 
absence) of tutorium; foveolate (versus not foveolate) 
prodorsum; lineolate and foveolate (versus not lineolate 
and foveolate) ventral body side.
E t y m o l o g y. The specific epithet ornamenta refers to the presence of striate--reticulate ornamentation 
on the notogaster in the new species.
Galumna (Galumna) villaclaraensis Ermilov sp. n.
(Figs 3–4)
T y p e  m a t e r i a l. Holotype (♀) and one paratype (♀): Cuba, Villa Clara Province, Baños de Elguea, 
soil litter with decaying leaves in a pasture under a tree 
(unknown data and collector; collection of the Tyumen 
State University Museum of Zoology, Tyumen, Russia).
The holotype is deposited in the collection of the 
Senckenberg Museum of Natural History, Görlitz, Germany; one paratype is deposited in the collection of the 
Tyumen State University Museum of Zoology, Tyumen, 
Russia. All specimens are preserved in 70% solution of 
ethanol with a drop of glycerol.
D i a g n o s i s  o f  a d u l t. Body length: 525–570. 
Rostrum concave medially. Lamellar and sublamellar 
lines divergent; L directed to insertion of rostral seta. 
Rostral seta long, setiform, slightly barbed; lamellar 
and interlamellar setae very long, subflagellate, slightly barbed; bothridial seta short, with clavate, broadly 
rounded distally, barbed head. Dorsosejugal porose area 
oval. Dorsosejugal suture complete. All notogastral setae represented by microsetae. Four pairs of porose areas rounded; la inserted posteriorly to area Aa. Median 
pore present, large. Opisthonotal gland located laterally to area A1; lyrifissure im close and anteriorly to area 
A1. Circumpedal carina short. Epimeral and anogenital 
Table 1. Leg setation and solenidia of adult Muliercula ornamenta Ermilov sp. n.
Leg
Tr
Fe
Ge
Ti
Ta
I
v′
d, (l), bv′′, v′′
(l), v′, σ
(l), (v), φ1, φ2 (ft), (tc), (it), (p), (u), (a), s, (pv), v′, (pl), l′′, ɛ, ω1, ω2
II
v′
d, (l), bv′′, v′′
(l), σ
(l), (v), φ
(ft), (tc), (it), (p), (u), (a), s, (pv), ω1, ω2
III
l ′, v′
d, l ′, ev′
l′, σ
l’, (v), φ
(ft), (tc), (it), (p), (u), (a), s, (pv)
IV
v′
d, ev′
d, l′
l’, (v), φ
ft′′, (tc), (p), (u), (a), s, (pv)
Notes. Roman letters refer to normal setae, Greek letters refer to solenidia (except ɛ – famulus); single quotation mark (′) designates 
setae on the anterior and double quotation (′′) setae on the posterior side of a given leg segment; parentheses indicate addition of both 
members of a pseudosymmetrical pair.

ERMILOV, TOLSTIKOV
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
Том 103
№ 10
2024
setae setiform, roughened. Postanal porose area oval. 
Leg setae l'' and v' absent on tarsus I; solenidion on tibia 
IV inserted in anterior part of the segment.
D e s c r i p t i o n  o f  a d u l t. Measurements. Body 
length: 525 (holotype), 570 (paratype); body width: 390 
(holotype), 375 (paratype).
Integument. Body color light brown. Body surface 
densely microfoveolate (visible under high magnification, ×1000); pteromorph partially with slight short striae; antiaxial side of leg femora I–IV and trochanters III 
and IV partially microtuberculate.
Prodorsum. Rostrum with semi-oval indentation 
(not visible in dorsal view versus clearly visible in 
dorsoanterior and anterior views). Lamellar line slightly 
thickened, directed to insertion of rostral seta but slightly not reaching it; sublamellar line thin, curving backwards. Rostral seta (90–94) setiform, slightly barbed; 
lamellar (154–165) and interlamellar (210–221) setae 
subflagellate, slightly barbed; bothridial seta (45–49) 
with short stalk and clavate, broadly rounded distally, 
barbed head; exobothridial seta represented by alveolus. 
Dorsosejugal porose area (11 × 7) oval, transversely oriented, located posteriorly to insertion of interlamellar 
seta. Dorsophragma medium--sized.
Notogaster. Dorsosejugal suture complete, but median part slightly developed. All notogastral setae 
Fig. 3. Galumna (Galumna) villaclaraensis Ermilov sp. n., adult (gnathosoma and legs not shown): a – dorsal view, b – ventral 
view, c – right lateral view, d – posterior view. Scale bar 100 µm.

	
NEW SPECIES AND NEW RECORDS OF ORIBATID MITES (ACARI, ORIBATIDA)
9
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ
Том 103
№ 10
2024
represented by microsetae (2). Four pairs of porose areas (15–19) rounded, clearly bordered; la inserted posteriorly to area Aa. Median pore present, single, comparatively large (11). Opisthonotal gland opening and all 
lyrifissures distinct; gla located laterally to area A1; im 
close and anteriorly to area A1.
Gnathosoma. Generally, similar to Galumna (e. g. Ermilov et al., 2023). Size of subcapitulum: 131–139 × 
101–105; subcapitular setae (a, h: 26–30; m: 19–22) setiform, slightly barbed; m thinnest; both adoral setae (17–
19) setiform, barbed. Length of chelicera: 142–146; cheliceral setae (cha: 60; chb: 37–41) setiform, barbed. Length 
of palp: 79–82; postpalpal seta (7) spiniform, smooth.
Epimeral region. Anterior margin of ventral plate 
smooth. Epimeral setal formula: 1–0–1–2. Setae 
(1a, 3b: 45–49; 4a, 4b: 30–37) setiform, roughened. 
Fig. 4. Galumna (Galumna) villaclaraensis Ermilov sp. n., adult: a – rostrum, anterior view; b – subcapitulum, ventral view; 
c – leg I (without trochanter), right, antiaxial view; d – leg II (without tarsus and trochanter), right, antiaxial view; e – leg 
III (without tarsus), left, antiaxial view; f – leg IV, left, antiaxial view. Scale bars 50 µm.